Evening Primrose

Oenothera is a genus of about 125 species of annual, biennial and perennial herbaceous flowering plants, native to North and South America. It is the type genus of the family Onagraceae, the Evening Primrose Family. Common names include evening-primrose, suncups, and sundrops.
The species vary in size from small alpine plants 10 cm tall (e.g. O. acaulis from Chile), to vigorous lowland species growing to 3 m (e.g. O. stubbei from Mexico). The leaves form a basal rosette at ground level and spiral up to the flowering stems; the leaves are dentate or deeply lobed (pinnatifid). The flowers open in the evening, hence the name "evening-primrose", and are yellow in most species but white, purple, pink or red in a few. Most native desert species are white. The fragrant tufted evening-primrose Oenothera caespitosa, a Southwestern species, first blooms white but turns pink or light magenta.
One of the most distinctive features of the flower is the stigma with four branches, forming an X shape. Pollination is by Lepidoptera (moths) and bees; like many members of the Onagraceae, however, the pollen grains are loosely held together by viscin threads (see photo below), meaning that only bees that are morphologically specialized to gather this pollen can effectively pollinate the flowers (it cannot be held effectively in a typical bee scopa). Furthermore, the flowers are open at a time when most bee species are inactive, so the bees which visit Oenothera are also compelled to be vespertine temporal specialists. The seeds ripen from late summer to fall.